- I am very sad about what happened at the zoo, not least because Santino was killed even though he was old and hardly dangerous, says world-renowned primatologist Frans de Waal at Emory University in Atlanta to SVT.
Santino showed us that chimpanzees have abilities that we previously thought only humans had.
One behavior that fascinated the researchers was that he collected and hid rocks before the park opened.
He would then throw the stones at visitors.
The staff has emptied hundreds of stone caches over the years.
- The discovery is remarkable for Santino planned spontaneously for the future.
Because he acted on his own, the result was more convincing than any experiment, says primatologist and professor Frans de Waal.
See the three biggest discoveries thanks to the chimpanzees in Furuvik in the clip.